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Header Images: Opilionid by Joe Warfel, Lynx Spider by Brian Reynolds, Lasiochernes cretonatus Pseudoscorpion by Anonymous.

Journal of Arachnology (JoA)

The Journal of Arachnology (ISSN 0160-8202), is a publication devoted to the study of Arachnida. It is published three times each year by the American Arachnological Society. Subscriptions accompany society membership or may be purchased by institutions. Learn about Membership here.

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Instructions to Authors

Revised January 2018

pdf version

All manuscripts are now submitted online at
General: The Journal of Arachnology publishes scientific articles reporting novel and significant observations and data regarding any aspect of the biology of arachnid groups. Articles must be scientifically rigorous and report substantially new information. Submissions that are overly narrow in focus (e.g., local faunal lists, descriptions of a second sex or of a single species without additional discussion of the significance of this information), that have poorly substantiated observational data, or that present no new information will not be considered. Book reviews will not be published.
Manuscripts must be in English and should use the active voice throughout. Authors should consult a recent issue of the Journal of Arachnology for additional points of style. Manuscripts longer than three printed journal pages (12 or more double-spaced manuscript pages) should be prepared as Feature Articles, shorter papers as Short Communications. Invited Reviews will be published from time to time and unsolicited reviews are also welcomed. All reviews will be subject to the same review process as other submissions.

Submission: Manuscripts should be prepared in Microsoft Word and then submitted electronically via our online system, PeerTrack (http://www.editorialmanager.com/arachno). PeerTrack will guide you through the step-by-step process including uploading the manuscript and all of its parts. The paper can be uploaded as one piece, with tables, figures, and appendices embedded, or as text, then tables, figures, and appendices, each uploaded individually. Ultimately, PeerTrack will assemble all parts of the paper into a PDF that you, as corresponding author, will need to approve before the submission process can be completed. Supplemental Materials (see below) can also be uploaded, but they are not bundled into the PDF.

Voucher Specimens: Specimens of species used in your research should be deposited in a recognized scientific institution. All type material must be deposited in a recognized collection/institution and the identity of the collection must be given in the text of the manuscript.
Checklist—Common Formatting Errors is available as a PDF at
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All Articles
All articles should be left-justified (not left and right justified) and should have page numbers. Accepted revised manuscripts should have line numbers.

Featured Articles
Title page.—The title page includes the complete name, address, and e-mail address of the corresponding author; the title in bold text and sentence case; each author’s name and address; and the running head.

Running head.—This should be in all capital letters, not exceeding 60 characters and spaces, and placed at the top of the title page. It should be composed of the authors’ surnames and a short title. Examples: SMITH—SALTICIDS OF PANAMA; SMITH & CRUZ—SALTICIDS… ; SMITH ET AL.—SALTICIDS…

Abstract.—Length: ≤ 250 words for Feature Articles; ≤ 150 words for Short Communications.

Keywords.—Give 3–5 appropriate keywords or phrases following the abstract. Keywords should not duplicate words in the title.

Text.—Double-space text, tables, legends, etc. throughout. Except for titles and headers, all text should be left-justified.  Do not add line numbers—they are automatically added by PeerTrack. Three levels of heads are used.

• The first level (METHODS, RESULTS, etc.) is typed in capitals and centered on a separate line. 
• The second level head begins a paragraph with an indent, is in bold type, and is separated from the text by a period and a dash. 
• The third level may or may not begin a paragraph but is italicized and separated from the text by a colon.

Use only the metric system unless quoting text or referencing collection data. If English measurements are used when referencing collection data, then metric equivalents should also be included parenthetically. All decimal fractions are indicated by a period (e.g., 3.141). Include geographic coordinates for collecting locales if possible, using one of the following formats: 0°12’32”S, 29°52’17”E or 0.2089°S, 29.8714°E.

Citation of references in the text: Cite only papers already published or in press. Include within parentheses the surname of the author followed by the date of publication. A comma separates multiple citations by the same author(s) and a semicolon separates citations by different authors, e.g., (Smith 1970), (Jones 1988; Smith 1993), (Smith & Jones 1986, 1987; Jones et al. 1989). Include a letter of permission from any person who is cited as providing unpublished data in the form of a personal communication.

Citation of taxa in the text: Include the complete taxonomic citation (author, year) for each arachnid genus and/or species name when it first appears in the abstract and text proper. For example, Araneus diadematus Clerck, 1757. For Araneae, this information can be found online at www.wsc.nmbe.ch. Citations for scorpions can be found in the Catalog of the Scorpions of the World (1758–1998) by V. Fet, W.D. Sissom, G. Lowe & M.E. Braunwalder. Citations for the smaller arachnid orders (pseudoscorpions, solifuges, whip scorpions, whip spiders, schizomids, ricinuleids and palpigrades) can be found at museum.wa.gov.au/catalogues-beta/. Citations for some species of Opiliones can be found in the Annotated Catalogue of the Laniatores of the New World (Arachnida,Opiliones) by A.B. Kury.

Literature cited.—Use the following style and formatting exactly as illustrated; include the full unabbreviated journal title. Personal web pages should not be included in Literature Cited. These can be cited within the text as (John Doe, pers. website) without the URL. Institutional websites may be included in Literature Cited. If a citation includes more than six authors, list the first six and add "et al." to represent the others.

citation format

Footnotes.—Footnotes are permitted on the first page, only to give current address or other author information, and at the bottom of tables (see below).

Taxonomic articles.—Consult a recent taxonomic article in the Journal of Arachnology for style or contact a Subject Editor for Systematics. Papers containing original descriptions of focal arachnid taxa should be listed in the Literature Cited section.

Tables.—Each table, with the legend above, should be placed on a separate manuscript page. Only horizontal lines (usually no more than three) should be included. When necessary, tables may have footnotes, for example, to specify the meanings of symbols about particular data.

Illustrations.—Original illustrations include photographs, line drawings, maps, and other graphic representations. All should be considered figures and numbered consecutively with other figures. You should ensure that all illustrations, at submission, are at high enough resolution to be useful to editors and reviewers; 300 dpi is usually sufficient. 
At the discretion of the Editor-in-Chief, a figure can be rendered in color in the online version but in monochrome in the journal’s printed version, or in color in both versions if warranted by the figure's context and content. Most figures will be reduced to single-column width (9 cm, 3.5 inches), but large plates can be printed up to two-columns width (18 cm, 7 inches). Address all questions concerning illustrations to the Editor-in-Chief of the Journal of ArachnologyDeborah Smith, Editor-in-Chief [E-mail: debsmith@ku.edu].
Legends for illustrations should be placed together on the same page(s). Each plate must have only one legend, as indicated below:

Figures 1–4. A-us x-us, male from Timbuktu.: 1. Left leg. 2. Right chelicera. 3. Dorsal aspect of genitalia. 4. Ventral aspect of abdomen.

The following alternate Figure numbering is also acceptable:

Figure 1a–e. A-us x-us, male from Timbuktu: a. Left leg. b. Right chelicera. c. Dorsal aspect of genitalia. d. Ventral aspect of abdomen.

Assemble manuscript.—The manuscript should be assembled in the following sequence: title page, abstract, text, tables with legends, figure legends, figures. As noted above, at the time of submission the paper can be uploaded as one piece, with tables, figures, and appendices embedded, or as text, then tables, figures, and appendices, each uploaded individually.

Supplemental materials.—Authors may submit for online publication materials that importantly augment the contents of a manuscript. These may be audio files (e.g., .mp3, .m4a, .aif, .wav), video files (e.g., .mov, .m4v, .flv, .avi), or Word documents (e.g., .doc, .docx) for large tables of data. Consult with the Editor-in-Chief if you are considering submitting other kinds of files. Audio and video files should be carefully edited before submission to eliminate leaders, trailers, and other extraneous content. Individual files may not exceed 10MB; no more than five files may be included as supplemental materials for a manuscript.

Supplemental materials will be considered by reviewers and therefore must be included at the time of manuscript submission. Supplemental materials are published online at the discretion of the editors.


Short Communications

Short Communications are usually limited to 3–4 journal pages, including tables and figures (11 or fewer double-spaced manuscript pages including Literature Cited; no more than 2 figures or tables). Internal headings (METHODS, RESULTS, etc.) are omitted. Short communications must include an abstract and keywords.

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Page charges.—Page charges are voluntary, but authors who are not members of the American Arachnological Society are strongly encouraged to pay in full or in part for their articles ($75 per journal page).

Proofs.—The Journal’s expectation is that the final revision of a manuscript, the one that is ultimately accepted for publication, will not require substantive changes. Accordingly, the corresponding author will be charged for excessive numbers of changes made in the proofs.

Reprints.—Hard copy reprints are available only from Allen Press via EzReprint, a user-friendly, automated online system for purchasing article reprints. If your paper is accepted, prior to its publication you will receive an e-mail containing both a unique URL (SmartLink) from Allen Press/Yurchak Press and information about the reprint order process. Clicking on the SmartLink will take you directly to a web portal where you may place your reprint order. The email will be sent to you from:  reprints@authorbilling.com. PDFs of papers published in theJournal of Arachnology are available to AAS members at the society’s web site. They are also available through BioOne (www.bioone.org) and JSTOR (www.jstor.org) if you or your institution is a member of BioOne or JSTOR. PDFs of articles older than one year are freely available from the AAS website.

Cover Artwork

Authors are encouraged to send high quality color photographs to the Editor-in-Chief to be considered for use on the cover. Images should be at least 300 dpi.

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Editors and Editorial Board

Editor-in-Chief: Deborah Smith
Department of Ecology & Evolutionary Biology, University of Kansas
Haworth Hall, 1200 Sunnyside Ave., Lawrence, KS 66045, USA 

Managing Editor: Richard S. Vetter
Department of Entomology 
University of California, Riverside
Riverside, CA USA 92521
E-mail: rick.vetter@ucr.edu

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Associate/Subject Editors

Systematics (Araneae) Subject Editor: Michael Rix 
Queensland Museum
Grey St. P.O. Box 3300 
South Brisbane, Queensland 4101, AUSTRALIA 

Systematics (non-Araneae) Subject Editor: Mark Harvey
Senior Curator, Department of Terrestrial Invertebrates, Western Australian Museum 
Francis Street, Perth, Western Australia 6000, AUSTRALIA
E-mail: mark.harvey@museum.wa.gov.au

Morphology and Physiology Subject Editor : Peter Michalik
Zoologisches Institut und Museum, Greifswald
Johann Sebastian Bach St. 11/12
E-mail: michalik@uni-greifswald.de

Behavior Subject EditorThomas C. Jones
Department of Biological Sciences
East Tennessee State University
Johnson City, TN 37614

Ecology Subject Editor: Martin Entling
Institute for Environmental Sciences, University of Koblenz-Landau 
Fortstr. 11, 76829 Landau, Germany

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Editorial Board

Alan Cady -- Miami University (Ohio)
Jonathan A. Coddington -- Smithsonian Institution
William Eberhard -- Universidad de Costa Rica
Rosemary Gillespie - University of California, Berkeley
Charles Griswold -- California Academy of Sciences
Marshal Hedin -- San Diego State University 
Marie Herberstein - Macquarire University
Yael Lubin - Ben-Gurion University of the Negev
Brent Opell -- Virginia Polytechnic Institute & State University
Ann L. Rypstra -- Miami University, Ohio
William A. Shear-- Hampden-Sydney College
Jeffrey Shultz - University of Maryland 
Petra Sierwald -- Field Museum 
Soren Toft - Aarhus University
I-Min Tso - Tunghai University (Taiwan)

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Undelivered and Back Issues

Contact for Undelivered Issues: Brian Patrick
Department of Biological Sciences, Dakota Wesleyan University
Mitchell, SD 57301 USA. 
Email: brpatric@dwu.edu

Contact for Back issues: Jim Carrel - download a pdf of presently available issues here
209 Tucker Hall 
University of Missouri - Columbia 
Columbia MO 65211 USA 
E-mail: carrelj@missouri.edu back to top

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Immediate Web Access to Your Accepted JoA Articles

Once your article is accepted, you will have the opportunity to purchase immediate open access to it on the AAS website. This means that anyone will be able to download your article as a pdf from the AAS website as soon as it is published there (usually just before the paper issues are mailed). Normally, JoA articles are embargoed for a year. For a reasonable price, you can make your article more available and help the AAS defray costs the society pays for on-line publication..

For paying by personal check:

  1. Download a pdf of the payment form and send it to the AAS treasurer: Cara Shillington (2017-2019), Biology Department, Eastern Michigan State University, Ypsilanti, MI 48197 USA.
  2. Please send an email to the AAS website administrator, Daniel Gloor webmaster@americanarachnology.org, alerting him that you have chosen this option. Include first author's name and the title of the article.

For paying on-line:

  1. Use the on-line order option below to pay for immediate access.
  2. Please send an email to the AAS website administrator, Daniel Gloor webmaster@americanarachnology.org, alerting him that you have chosen this option. Include first author's name and the title of the article.

Article Type


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Feature Article


Short Communication



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