Graduate Studies in Arachnology:
Behavior, Behavioral Ecology, and Neurophysiology


Updated August 14, 2008

 

List of Graduate Studies Sponsors
-- Links to Further Details

R. Jackson 
E. Jakob 
Y. Lubin
S. Masta
D. Morse
L. Rayor
S. Riechert

 


MAYDIANNE C.B. ANDRADE, Assistant Professor
University of Toronto at Scarborough

Degrees offered: M.Sc., Ph.D.

Address: Division of Life Sciences
University of Toronto at Scarborough
1265 Military Trail
Scarborough, ON M1C 1A4
Canada

Voice: (607) 287-7425 (office) and (607) 287-7124 (lab); FAX: (607) 287-7642

e-mail: mandrade@scar.utoronto.ca

Professional website: www.scar.utoronto.ca/~mandrade

Research Interests and Information on Graduate Program: Behavioural Ecology and Evolution of mating systems in spiders and insects.
Particular interests: Sexual selection, male mating strategies, sexual cannibalism and male self-sacrifice in spiders. Current research concentrates on Latrodectus spiders, but lab members are free to choose their own projects. Research includes field and lab studies directed at answering evolutionary and functional questions.
Students interested in mechanisms of behaviour (particularly if relevant to sperm competition or courtship signalling) are also welcome to apply. Students would be members of the Department of Zoology, which is a 3-campus-wide department (U Toronto, UT Mississauga, UT Scarborough).
Teaching assistantships, some internal fellowships, and other support are available for graduate students; however, students are encouraged to apply for external fellowships wherever possible.

updated 11 - 27 -2000

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FRIEDRICH G. BARTH, O. Univ.-Prof. Dr
Biozentrum, Institut für Zoologie, Universität Wien

Degrees offered: MSc (Diplom) and PhD (Dr.rer.nat.)

Address: Biozentrum
Institut für Zoologie
Universität Wien
Althanstraße14
A-1090 Wien
Austria

Voice: 0043/1/31336/1205; FAX: 0043/31336/778;

e-mail: Friedrich.G.Barth@univie.ac.at

Research Interests and Information on Graduate Program: Sensory systems (mechano/photo/chemo/thermo & hygro). Neuroethology. Biomechanics. Vibratory communication. Spider webs. Communication in stingless bees. Additional information on request.

updated 1 - 14 -2000
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TODD BLACKLEDGE, Assistant Professor
University of Akron

Degrees offered: Master of Science and Ph.D.

Address: Department of Biology
University of Akron
Akron, OH 44325-3908
USA

Voice: (330) 972-7264; FAX: (330) 972-8445
e-mail: blackledge@uakron.edu

Website: http://www3.uakron.edu/biology/

Research Interests and Information on Graduate Program: My research focuses upon the interface of animal behavior with ecological and evolutionary processes. Web weaving spiders provide a unique opportunity to integrate behavioral research across levels of biological organization for several reasons. Unlike the relatively ephemeral foraging and defensive behaviors of most animals, spider webs are quantifiable records of spiders’ behavioral decisions that endure for the life of webs. Also, the architectures of spider webs can be easily manipulated, allowing direct investigation of the ecological consequences of the behavioral decisions made by spiders during web construction. Furthermore, the function of webs depends upon the physical properties of the silks used to construct them, which are themselves dependent upon protein structure. Therefore, behavioral and genotypic evolution can be tightly linked in this system. My laboratory is set up to support a variety of research.

I have three main lines of current research:

  • behavioral ecology of orb web function
  • evolution of web architectures
  • biomechanical function of silk.

The Department of Biology at University of Akron emphasizes integrative research and has strong programs in both evolutionary ecology and physiological ecology. The department has grown rapidly over the last ten years to include many energetic new faculty, a new field station, and a strong research emphasis. The department offers a Master of Science as well as a PhD through a joint program with Kent State University.

added 1 - 15 - 2005

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JAMES CARREL, Professor
Division of Biological Sciences, Missouri University

Degrees offered: M.A./Ph.D.

Address: Division of Biological Sciences
Missouri University
105 Tucker Hall
Columbia, MO 65211 USA

Voice: (314) 882-0123;

e-mail: carrel@biosci.mbp.missouri.edu

Research Interests and Information on Graduate Program: Physiology, behavior, and ecology of spiders. Pheromones in spiders. Conservation biology of rare spiders endemic to scrub habitats in Florida. Teaching assistantships and special scholarships for gifted or underrepresented students are available to qualified candidates.
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GARY DODSON, Professor of Biology
Biology Department, Ball State Univ.

Degrees offered: M.S./Ed.D.

Address: Biology Department
Ball State University
Muncie, IN 47306 USA

Voice: (765) 285-8859; FAX: (765) 285-8804

e-mail: gdodson@bsu.edu

Professional website: http://www.bsu.edu/web/biology/faculty/dodson.html

Research Interests and Information on Graduate Program: Behavioral ecology of spiders, especially mating systems, male fighting behavior of thomisids. Also occurrence and significance of nectar feeding in spiders. Teaching assistantships are available to qualified candidates.

updated 1 - 19 - 2005
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DAN FORMANOWICZ, Professor and Graduate Advisor
The University of Texas at Arlington

Degrees offered: M.S.; Ph.D.

Address: Dept. of Biology
Box 19498
The University of Texas at Arlington
Arlington, TX 76019 USA

Office: (817) 272-2422; FAX: (817) 272-2855

e-mail: formanowicz@uta.edu

Professional website: www.uta.edu/biology/faculty/formanowicz/index.html

Departmental website: www.uta.edu/biology

Research Interests and Information on Graduate Program: Behavioral ecology of scorpions and spiders including; predator-prey interactions (foraging behavior, defensive behavior); temporal and geographic life history variation; costs of reproduction; costs and benefits of burrowing. There are currently 5 Ph.D. and 5 M.S. students in the lab. working on a variety of questions within the general areas listed above. The research in our laboratory also focuses on reptile and amphibian systems. Information about the graduate programs at UTA can be found on the department website and the UTA departmental graduate program website.

Recent Publications:

Shaffer, L.R. & D.R. Formanowicz, Jr. 2000. Sprint speeds of juvenile scorpions: among family differences and parent-offspring correlations. J. Insect Behavior 13:45-54.

Amaya, C.C., P.D. Klawinski, & D.R. Formanowicz, Jr. The effects of leg autotomy on running speed and foraging ability in two species of wolf spider. Am. Midl. Nat., in press.

updated 6 - 26 -2000
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EILEEN A. HEBETS, Assistant Professor
School of Biological Sciences, University of Nebraska, Lincoln

Degrees offered: Ph. D

Address: School of Biological Sciences
University of Nebraska
Lincoln, NE 68588 USA

Office Phone: (402) 472-2571

E-mail: ehebets@unlnotes.unl.edu

Professional website: http://www.biosci.unl.edu/faculty/FacPages/hebets.asp

Research Interests and Information on Graduate Program: I am interested in the evolution of complex signaling, sexual selection, and learning and memory in arachnids. My research programs are broad encompassing wolf spiders, jumping spiders, and amblypygids.

new listing October 27, 2005

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ROBERT R. JACKSON, Associate Professor
Department of Zoology, University of Canterbury

Degrees offered: M.Sc./Ph.D.

Address: Department of Zoology
University of Canterbury
Private Bag 4800
Christchurch New Zealand

Voice: +64 3 364-2064 [or] +64 3 364-2071; FAX: +64 3 364-2024;

e-mail: r.jackson@zool.canterbury.ac.nz

Research Interests and Information on Graduate Program: Aggressive mimicry.Foreign students are subject to fees. Financial support from within New Zealand for foreign students is virtually non-existent.
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ELIZABETH M. JAKOB, Associate Professor
University of Massachusetts, Amherst

Degrees offered: M.S./Ph.D.

Address: Department of Psychology
Tobin Hall
University of Massachusetts
Amherst, MA 01003 USA

Voice: (413) 577-0707; FAX: (413) 545-0231

e-mail: ejakob@ent.umass.edu

Professional website: http://www.umass.edu/ent/DeptSite/jakob.html

Research Interests and Information on Graduate Program: Behavior of spiders, including group living in pholcids, and learning and the effects of environmental enrichment in salticids. I am also working a project to evaluate several salticid species for their efficacy as predators in greenhouses. I would be interested in supervising students on other projects as well.

I can sponsor students in the Department of Entomology, the Program in Organismic and Evolutionary Biology, and the Program in Neuroscience and Behavior. These differ substantially in structure, so please look at the website descriptions to see which fits you best. Some teaching assistantships are available, and research assistantships are occasionally available as well. Please contact me by email if you are interested in UMass.

last updated 1 - 13 - 2000
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YAEL D. LUBIN, Associate Prof.
Department of Life Sciences, Ben Gurion Univ.

Degrees offered: M.Sc./Ph.D.

Address: Mitrani Center for Desert Ecology
Blaustein Institute for Desert Research
Ben Gurion Univ.
Sede Boker Campus, 84993 ISRAEL

Voice: ++972-7-552650, [or] ++972-7-565073; FAX: ++972-7- 557829;

e-mail: lubin@bgumail.bgu.ac.il

Research Interests and Information on Graduate Program: Behavioural ecology, evolution of sociality, ecology. Some current research activities (including work of students and post-doctoral fellows):

  • Social organization in spiders and preconditions for sociality. I am working on Stegodyphus (Eresidae), including two African species which are highly social and a species found in the semi-arid parts of Israel which is subsocial. I am particularly interested in the conditions influencing dispersal (and reduced dispersal) in these species.
  • Mate selection and consequences of sexual competition and female choice. This work is currently on two species, Stegodyphyus lineatus (Eresidae) and Latrodectus revivensis (Theridiidae).
  • Genetical and environmental influences on behaviour. This is a study of a species of agelenid which demonstrates different egg-sac production strategies that vary in their frequency of occurrence in different populations.
  • The influence of vegetation structure on web-building in spiders, a study at two levels, the individual and the community.
  • Foraging behaviour of spiders. A variety of subjects are included here, e.g., prey capture behaviour of specialized spider predators (gnaphosids, salticids), foraging decisions of S. lineatus.

Financial support: Some support is available for MSc and PhD students from the Dept. of Life Sciences, the Blaustein International Center and the Mitrani Center for Desert Research. Post-doctoral fellowships may be available through the Blaustein International Center and form grants. Students may register for degrees at Ben Gurion University, or may come here from other universities to do their field work at the Mitrani Center (in which case they usually have support from their 'home' university). MSc students must take courses at the main campus in Beer Sheva. PhD students do not have course requirements. A variety of desert and steppe habitats are easily accessible here. Field studies are encouraged (lab space is limited). Students live on the Sede Boker Campus, 50 km south of the town of Beer Sheva.

Further information is available upon request.
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SUSAN MASTA, Assistant Professor
Department of Biology, Portland State University

Degrees offered: M.S./ Ph.D.

Address: Department of Biology
Portland State University
P.O. Box 751
Portland, OR 97207 USA

Voice: (503) 725-8505 (office); FAX: (503) 725-3888

e-mail: smasta@pdx.edu

Professional website: http://web.pdx.edu/~smasta/

Departmental website: http://www.orgbio.pdx.edu/

Research Interests and Information on Graduate Program: Biological diversification, from the molecular to organismal levels, with a focus on diversification of arachnid groups and divergence among jumping spider populations. I have specific interests in RNA and genome evolution, population genetics, phylogenetics, and sexual selection. Information on the graduate program is available on the departmental website. Departmental teaching assistantships are available, with the possibility of research assistantships.

Added June 22, 2005
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DOUGLASS H. MORSE, Professor
Department of Ecology and Evolutionary Biology, Brown University

Degree offered: Ph.D.

Address: Department of Ecology and Evolutionary Biology
Box G-W
Brown University
Providence, RI 02912 USA

Voice: (401) 863-3152; FAX: (401) 863-2166;

e-mail: d_morse@brown.edu

Professional website: http://www.brown.edu/Departments/EEB/faculty/Morse.html

Research Interests and Information on Graduate Program: Ecology and behavior of spiders. I currently focus on foraging, life histories and sexual selection, primarily with the crab spider Misumena vatia (Thomisidae). Recent work has included the role of experience on foraging behavior and patch choice at different stages of the life cycle; and the roles played by sex ratios, aggression, mate-guarding, and mate-finding in determining male reproductive strategy. In a second, newer line of research, I study the movements and population dynamics of an intertidal wolf spider Pardosa lapidicina that moves up and down cobble beaches with the tides. Graduate students are supported with fellowships and assistantships.

updated 1 - 25 - 2000
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LINDA S. RAYOR, Assistant Professor
Dept of Entomology, Cornell University

Degrees offered: M.S./Ph.D.

Address: Department of Entomology
Cornell University
Ithaca, NY 14853 USA

Voice: (607) 255-8406
FAX: (607) 255-0939

e-mail: LSR1@cornell.edu

Websites: http://www.entomology.cornell.edu/Faculty_Staff/Rayor

Research Interests and Information on Graduate Program: Behavioral ecology, with an emphasis on social behavior, mother-offspring interactions, learning, and tradeoffs between foraging and predation risk. Current projects in my laboratory involve: (1) Social behavior in the atypical Australian huntsman spider, Delena cancerides and (2) Social behavior and tactile communication in amblypygids. I am actively seeking graduate students for either of these projects for Fall 2003. I would consider graduate students interested in other aspects of spider behavior. Teaching Assistantships are available for qualified applicants.

added 8 - 21 - 2002
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SUSAN E. RIECHERT, Professor
Dept. of Ecology & Evolutionary Biology, Univ. of Tennessee

Degree offered: Ph.D.

Address: Department of Ecology & Evolutionary Biology
University of Tennessee
Knoxville, TN 37996-0810 USA

Voice: (685) 974-2371; FAX: (685) 974-3067

e-mail: sriecher@utk.edu

Research Interests and Information on Graduate Program: Ecology, behavior, evolutionary biology, genetics, biocontrol. Prefer students with prior research experience (e.g., a masters degree).

last updated 1 - 14 - 2000
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J. ANDREW ROBERTS, Assistant Professor
Department of Evolution, Ecology, and Organismal Biology
Ohio State University, Newark

Degrees offered: M.S./Ph.D.

Address: Ohio State University, Newark
Department of Evolution, Ecology, and Organismal Biology
1179 University Drive
Newark, OH 43055 USA

Voice: (740) 366-9178


FAX:  (740) 366-5047

e-mail: roberts.762@osu.edu

Websites: Professional homepage: http://www.biosci.ohio-state.edu/~eeob/faculty/roberts.html

Departmental Homepage:  http://www.biosci.ohio-state.edu/~eeob/


Research Interests and Information on Graduate Program: At present, much of my research concerns constraints on the evolution of animal signals and signaling behavior.  My main research interests are in the evolution of animal communication and social behavior using spider model systems.  I apply an integrative approach to my research program, incorporating techniques from several disciplines (e.g. environmental measurements, field and laboratory observation and experimentation, digital imaging and video/audio playback, chemical extraction, DNA microsatellites) to address interesting questions about the evolution of animal signals, sexual selection and mate choice, and group formation and social behavior.  I have collaborated on a variety of projects working with invertebrate and vertebrate organisms in habitats including coastal riparian, flood plain, old growth forest, and cave and karst ecosystems.  Graduate students under my supervision will be enrolled in the Department of Evolution, Ecology, and Organismal Biology at the Ohio State University, Columbus Campus.

added 1 - 5 - 2005

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ANN L. RYPSTRA, Associate Prof. of Zoology
Department of Zoology, Miami University

Degrees offered: M.S./Ph.D.

Address: Department of Zoology
Miami University
Oxford, OH 45056 USA
(I am officially located on the Hamilton Campus of Miami so the address on my publications is different but it is more convenient if I hear from prospective students at this address);

FAX: (513) 529-6900;

e-mail: rypstral@MUOhio.edu

Research Interests and Information on Graduate Program: Spacing patterns in spiders, evolution of social behavior, spiders in agroecosystems, food web ecology, foraging behavior, influence of spiders on the foraging activity of herbivores. All students admitted receive a graduate assistantship. Most are required to teach. Program is a broad based zoology program but the departmental emphasis is ecology. Of 33 faculty, 15 conduct research in some aspect of ecology. Students have to have had undergraduate courses in ecology, physiology, genetics, statistics, one year of physics and two years of chemistry, but some of these courses can be taken after the student is admitted and comes to Miami. GREs are required.
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CARA SHILLINGTON Associate Professor
Dept of Biology, Eastern Michgan University

Degrees offered: M.Sc.

Address: 316 Mark Jefferson
Department of Biology
Eastern Michigan University
Ypsilanti, MI 48197 USA

Voice: (734) 487-4433 (office/lab) and (734) 487-4242 (department office)

e-mail: cara.shillington@emich.edu

Professional Homepage: http://www.emich.edu/biology/details.php?dep=Biology&ID=49

Research Interests and Information on Graduate Program: My research focuses on an integrative, field-based, evolutionary approach to study the behavioral and physiological ecology of tarantulas. Current research topics include intra- and interspecific variation in metabolic rates, life history variation, mating systems, and foraging behavior.
Teaching assistantships are available to most students accepted into our department. In addition, the biology department has substantial fellowships that are awarded annually.

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W. DAVID SISSOM, Professor
Dept. of Life, Earth, & Environmental Sciences, West Texas A&M University

Degrees offered: M.S.

Address: Dept. of Life, Earth, & Environmental Sciences
West Texas A&M University
WTAMU Box 60808
Canyon, TX 79016 USA

Voice: (806) 651-2578; Fax: (806) 651-2928

e-mail: dsissom@mail.wtamu.edu

Departmental website: http://www.wtamu.edu

Research Interests and Information on Graduate Program: My primary research interests are in scorpion systematics, phylogeny (based on morphological characters), life history, behavior, and ecology. I have secondary interests in systematics and regional inventories of other arachnids. A limited number of teaching assistantships are available for qualified applicants.

last updated 4 - 1 - 2002
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GEORGE W. UETZ, Professor of Biological Sciences
University of Cincinnati

Degrees offered: M.S./Ph.D.

Address: Department of Biological Sciences
Univ. of Cincinnati
P.O. Box 210006
Cincinnati, OH 45221-0006 USA

Phone (office): (513) 556-9752, (lab) (513) 556-9753, (other) (513) 556-9700; FAX: (513) 556-5299;

e-mail: George.Uetz@uc.edu

Professional homepages:

Departmental homepage: http://www.biology.uc.edu

Research Interests: The major focus of my research is the behavioral ecology of spiders. I am interested in the selective forces shaping the evolution of social behavior, and use both field-based observational studies and laboratory experimentation in two long-term research projects:

  • Visual communication, species recognition and sexual selection - using video imaging and playback techniques to investigate the essential elements of signalling behavior in courtship, which functions in reproductive isolation and mate choice in Schizocosa wolf spiders (Lycosidae). Current work is directed at understanding the role of multi-modal communication (visual and vibratory signals) as an indicator of male condition in female mate choice. We are using audio-visual playback in cue-combination and cue-conflict experiments, to tease apart the elements of a complex communication system.
  • Group foraging in colonial web-building spiders - this project is directed at understanding the cost-benefit fitness tradeoffs involved in group-living (foraging success, predation risk, aggression, silk costs) for colonial web-building spiders of the genus Metepeira (Araneidae), which are "an exception to the rule"- among most (typically asocial) spiders (some work is being done in collaboration with Dr. Beth Jakob of U. Mass. and Dr. Craig Hieber of Saint Anselm College).
  • Two additional areas of research are:

    • Influence of El Nino on aggregative behavior and genetic structure of Metepeira populations on the California coast.
    • Role of habitat structure and refuge augmentation on spider populations and biological control of pests in agriculture (in Collaboration with Dr. Alan Cady of Miami University, Oxford, OH)

    I will also support student research in other areas of arachnology/behavioral ecology.

    Information About the Graduate Program: My research is funded by grants from the National Science Foundation, and my lab is well-equipped to pursue many types of arachnological research. I currently sponsor two graduate students, and have room for one to two more next year.
    Our department offers competitive assistantship and fellowship support (including tuition remission scholarships)for graduate students. Our graduate program has been re-organized recently, and the areas of Ecology, Evolution and Behavior have been strengthened by the addition of several new faculty.

    updated 1 / 19 / 2000
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    FRITZ VOLLRATH
    Department of Zoology, University of Aarhus

    Degrees offered: M.Sc./Ph.D.

    Address: Department of Zoology
    Universitetsparken B135
    DK 8000 Aarhus C Denmark

    Voice: +45-89 42 27 65 (Sec: +45- 89 42 27 27); FAX: +45-86 12 51 75;

    e-mail: vollrath@aau.dk

    Research Interests and Information on Graduate Program: Orb spider ecology, behaviour, web engineering, silk biomechanics. Information upon request.

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    SEAN WALKER, Assistant Professor
    California State University Fullerton

    Degree offered: M.S.

    Address: Department of BIological Science
    California State University Fullerton
    Fullerton CA 92831 USA

    Phone (office): (714) 278-3610; Phone (lab): (714) 278-8204; FAX: (714) 278-3426

    e-mail: swalker@fullerton.edu

    Professional website (homepage): http://biology.fullerton.edu/swalker

    Research Interests and Information on Graduate Program: My primary research interests are in the cognitive, behavioral, and evolutionary ecology of terrestrial invertebrates (mainly Arachnids and Orthopterans). In particular, I am interested in animal decision making as it relates to foraging and reproduction (e.g., mate choice & reproductive investment) and have a keen interest in any project related to animal communication, the evolution of sex differences, and the evolution of alternative tactics and strategies. I have also worked on projects related to the estimation and comparison of terrestrial invertebrate biodiversity and pollination biology.

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    DAVID H. WISE, Professor of Entomology
    University of Kentucky

    Degrees offered: M.S., Ph.D.

    Address: Department of Entomology,
    S-225 Ag Sci Bldg -- North
    University of Kentucky
    Lexington, KY 40546-0091 USA

    Phone (office): (606) 257-4264; Phone (lab): (606) 257-4693; FAX: (606) 323-1120;

    e-mail: dhwise@pop.uky.edu

    Professional website (homepage): http://www.uky.edu/~dhwise/wise.html

    Research Interests and Information on Graduate Program: Behavioral, population and community ecology of spiders in food webs of forest-floor leaf litter and vegetable crops. Please visit my website for detailed, current information on my group's research program.

    last updated 1 / 18 / 2000
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